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 Etiology of sperm DNA damage
http://fertilitysolutions.com.au/sperm-dna-fragmentation/
  DNA damage during spermatogenesis
 Hundreds of DNA breaks to initiate meiotic crossover recombination
  Thousands of DNA breaks to facilitate histone to protamine transition
“long term” storage in epididymis
Figure from Verver et al., 2015
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